The current paper discusses the growing biological issue of "cryptic" species. For species with substantial genetic variety but little morphological variation, a cryptic species notion was recently put out. In order to identify three new species and illustrate their dual nature, we use special material from a fascinating group of nudibranch molluscs of the genus Trinchesia from European waters as a study in itself. Each species in that complex has stable morphological and molecular distinctions, but their overall similarity can be determined as a "cryptic" species complex. This species complex may therefore be referred to as "cryptic," "pseudocryptic," or "non-cryptic." We further link the species problem to epigenetics and provide evidence for a very high rate of speciation in this species complex. When a finer multilayer morphological and molecular scale is employed to uncover more accurately defined species, the "cryptic" addition to "species" is no longer essential because the degree of crypticity is a continuum. This is supported by the broad discussion of the available metazoan-wide data in the current study. Molecular and morphological approaches should be used to create a fine-scale multilevel taxonomic framework. This does not necessarily mean that purely molecular-based "cryptic" species will be a posteriori transformed into morphologically defined "pseudocryptic" ones. The current study's implications are significant for various disciplines, such as fine-scale biodiversity assessments and conservation biology.